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Many mail servers are configured to reject mail if it is received from a sending server with inconsistent DNS. This is because the majority of SPAM mail comes from malware on infected computers. As the compromised machines were not meant to be used as mail servers, DNS will not have been created on the IP the malware sends from. Therefore, to minimise the possibility of the recipient's mail server rejecting mail from your server out of hand, it's important to ensure that your DNS is consistent.
To lower the chance of your mail being rejected, three things need to be set up: